I also have the mysterious varying oal when reloading 9mm and 45acp with my 550.
On my machine, when all 4 stations have brass in them, the oal is longer . If I just put a one in the bullet seating station, it seats to my preset oal. One of the other stations , it appears, reaches the of the press first and prevents the bullet from setting all the way.
I have tried all ideas I found by searching this forum as well as others. All stations are adjusted per dillons instructions.